/* A spacious academic website: white, navy, and a restrained gold accent. */
:root {
  --paper: #fff; --ink: #112d50; --body: #415063; --muted: #627083;
  --line: #dce3ea; --accent: #285d88; --soft: #f3f6f9; --card: #fff;
  --gold: #b89955; --sans: 'Helvetica Neue', Helvetica, Arial, sans-serif;
  --display: var(--sans);
}
html[data-theme="dark"] {
  --paper: #101d2d; --ink: #eef4fc; --body: #cbd7e6; --muted: #a5b6ca;
  --line: #334359; --accent: #a8d0f2; --soft: #182b40; --card: #182b40;
}
* { box-sizing: border-box; }
html { font-size: 16px; scroll-behavior: smooth; }
body { margin: 0; background: var(--paper); color: var(--body); font: 17px/1.75 var(--sans); -webkit-font-smoothing: antialiased; }
a { color: var(--accent); text-decoration: none; }
a:hover { text-decoration: underline; text-underline-offset: 5px; }
a:focus-visible, button:focus-visible, input:focus-visible { outline: 3px solid var(--gold); outline-offset: 5px; }
button, input { font: inherit; }
button { cursor: pointer; }
[hidden] { display: none !important; }
img { max-width: 100%; }
h1, h2, h3 { font-family: var(--sans); color: var(--ink); line-height: 1.18; font-weight: 650; letter-spacing: -.035em; }
h1, h2 { font-family: var(--display); font-weight: 700; letter-spacing: -.025em; }
h1 { font-size: clamp(38px, 4.4vw, 60px); margin: 0 0 22px; }
h2 { font-size: 36px; margin: 48px 0 22px; }
h3 { font-size: 25px; margin: 0 0 16px; }
p { margin: 0 0 22px; }
ul, ol { padding-left: 24px; }
li { margin-bottom: 10px; }
strong { font-weight: 650; }
::selection { background: #d7e7f3; color: #092751; }
main { outline: none; }
.skip-link { position: fixed; top: -100px; left: 20px; background: var(--ink); color: var(--paper); padding: 10px 20px; z-index: 20; }
.skip-link:focus { top: 15px; }
.site-header { background: var(--paper); border-bottom: 1px solid var(--line); }
.header-inner { max-width: 1400px; padding: 35px 5%; min-height: 136px; display: flex; gap: 30px; align-items: center; margin: auto; }
.wordmark { display: flex; flex-direction: column; color: var(--ink); font-family: var(--display); font-size: 29px; font-weight: 700; letter-spacing: -.8px; line-height: 1.2; white-space: nowrap; }
.wordmark:hover { text-decoration: none; }
.wordmark-degree { font-size: 17px; font-weight: 400; letter-spacing: -.2px; }
.wordmark-subtitle { font-family: var(--sans); font-size: 10px; font-weight: 400; letter-spacing: .3px; margin-top: 10px; }
.primary-links { margin-left: auto; display: flex; align-items: center; gap: 26px; }
.primary-links a { font-size: 18px; font-weight: 600; color: var(--ink); white-space: nowrap; padding: 10px 0; position: relative; }
.primary-links a::after { content: ''; height: 2px; position: absolute; bottom: 0; left: 0; width: 0; background: var(--gold); transition: width .2s; }
.primary-links a:hover { text-decoration: none; }
.primary-links a:hover::after, .primary-links a[aria-current="page"]::after { width: 100%; }
.appearance-toggle { width: 44px; height: 44px; flex-shrink: 0; border: 0; border-radius: 50%; background: var(--soft); color: var(--ink); font-size: 22px; padding: 0; line-height: 1; }
.menu-toggle { display: none; }
.content-width { width: min(1120px, 88%); margin-inline: auto; }
.eyebrow { font-size: 13px; font-weight: 600; letter-spacing: .4px; color: var(--accent); margin-bottom: 16px; line-height: 1.6; }
.home-hero { background: #020817; color: #fff; }
.hero-inner { width: min(1240px, 92%); margin: auto; display: grid; grid-template-columns: .9fr 1.1fr; align-items: center; gap: 28px; padding: 12px 0 20px; }
.hero-media { margin: 0; min-width: 0; width: 100%; max-width: 530px; justify-self: end; }
.hero-media img { width: 100%; height: auto; display: block; }
.image-caption { color: #adbed1; padding-top: 13px; font-size: 10px; letter-spacing: .4px; line-height: 1.6; text-align: center; }
.hero-card { min-width: 0; padding: 36px 0 36px 18px; }
.hero-card .eyebrow { margin-bottom: 23px; color: #9ddaca; font-size: 10px; letter-spacing: 1.7px; }
.hero-card h1 { color: #fff; font-size: clamp(40px, 4.2vw, 58px); margin: 0 0 25px; letter-spacing: -.045em; white-space: nowrap; }
.hero-card h1 span { font-size: .48em; font-weight: 400; letter-spacing: -.02em; }
.hero-tagline { font-size: clamp(20px, 2.15vw, 29px); line-height: 1.55; font-weight: 400; color: #d7e5f3; margin-bottom: 32px; }
.hero-actions { display: flex; flex-wrap: wrap; align-items: center; gap: 24px; }
.hero-about { color: #e4ecf5; font-size: 13px; font-weight: 500; }
.hero-about span { margin-left: 10px; }
.hero-card .button-primary { background: #fff; color: #092751; border-color: #fff; }
.hero-card .button-primary:hover { background: #c7eee5; border-color: #c7eee5; }
.button { display: inline-flex; justify-content: center; align-items: center; gap: 22px; min-height: 48px; font-size: 13px; font-weight: 600; padding: 13px 22px; border: 1px solid transparent; line-height: 1.5; border-radius: 0; }
.button-primary { background: var(--ink); color: var(--paper); border-color: var(--ink); }
.button-primary:hover { background: var(--accent); color: var(--paper); border-color: var(--accent); text-decoration: none; }
.button-row { display: flex; gap: 20px; flex-wrap: wrap; margin: 26px 0; }
.text-link { display: inline-flex; align-items: center; gap: 15px; font-size: 14px; font-weight: 600; color: var(--ink); }
.home-intro { display: grid; grid-template-columns: minmax(230px, 320px) minmax(0, 1fr); gap: 70px; align-items: start; padding-top: 66px; padding-bottom: 90px; scroll-margin-top: 30px; }
.home-intro h2 { margin: 0 0 26px; font-size: 38px; }
.profile-portrait { margin: 0; }
.profile-portrait img { display: block; width: 100%; height: auto; background: #fff; }
.profile-portrait figcaption { padding-top: 18px; border-top: 1px solid var(--line); margin-top: 18px; color: var(--ink); font-size: 16px; font-weight: 600; line-height: 1.6; }
.profile-portrait figcaption span { display: block; margin-top: 5px; color: var(--muted); font-size: 13px; font-weight: 400; }
.intro-copy p { font-size: 17px; line-height: 1.8; }
.intro-copy .eyebrow { font-size: 11px; line-height: 1.6; }
.intro-copy p:last-of-type { margin-bottom: 25px; }
.profile-links { display: flex; gap: 30px; flex-wrap: wrap; }
.vision-link { font-size: 14px; margin: 32px 0 0; padding-top: 24px; border-top: 1px solid var(--line); }
.home-research { padding: 74px 0 66px; background: var(--soft); }
.section-heading { display: flex; align-items: end; justify-content: space-between; gap: 30px; margin-bottom: 34px; }
.section-heading h2 { margin: 0; }
.section-heading .eyebrow { margin-bottom: 15px; }
.research-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 42px; }
.research-overview { border-top: 1px solid #a9b9ca; padding-top: 23px; color: var(--body); position: relative; display: block; }
.research-overview:hover { text-decoration: none; }
.research-overview:hover h3 { color: var(--accent); }
.direction-number { display: block; font-size: 12px; color: var(--accent); letter-spacing: 1.7px; margin-bottom: 22px; }
.research-overview h3 { font-size: 24px; line-height: 1.3; margin-bottom: 16px; }
.research-overview p { font-size: 15px; line-height: 1.8; margin-bottom: 12px; }
.research-arrow { font-size: 25px; color: var(--accent); }
.home-news { padding-top: 78px; padding-bottom: 90px; }
.update-row { display: grid; grid-template-columns: 125px 1fr; gap: 32px; border-top: 1px solid var(--line); padding: 24px 0; }
.update-row:last-child { border-bottom: 1px solid var(--line); }
.update-row time { font-size: 12px; color: var(--muted); letter-spacing: 1.2px; padding-top: 3px; }
.update-row p { margin: 0; font-size: 16px; }
.page-banner { background: var(--soft); border-bottom: 1px solid var(--line); }
.page-banner-inner { max-width: 1120px; width: 88%; margin: auto; padding: 66px 0 60px; }
.page-banner h1 { max-width: 1050px; }
.page-banner .eyebrow { margin-bottom: 16px; }
.section-intro { font-size: 21px; line-height: 1.65; max-width: 800px; margin: 0; }
.main-content { max-width: 1000px; width: 88%; margin: 0 auto; padding: 54px 0 90px; }
.main-content > p, .main-content > ul, .main-content > ol, .research-direction > p { max-width: 860px; }
.research-direction { border-top: 1px solid var(--line); margin-top: 38px; padding-top: 30px; padding-bottom: 10px; scroll-margin-top: 30px; }
.research-direction::after { content: ''; display: block; clear: both; }
.research-illustration { float: right; width: 36%; margin: 5px 0 22px 36px; }
.research-illustration img { display: block; width: 100%; height: auto; }
.research-illustration figcaption { font-size: 12px; color: var(--muted); line-height: 1.7; margin-top: 12px; }
.research-direction h2 { margin-top: 0; font-size: 31px; scroll-margin-top: 25px; }
.research-direction .direction-number { margin-bottom: 16px; }
.research-jump-links { display: flex; flex-wrap: wrap; gap: 12px 24px; margin: 28px 0 10px; font-size: 13px; }
.research-jump-links a { padding: 6px 0; }
.home-applications { padding-block: 78px 0; scroll-margin-top: 30px; }
.applications-intro { max-width: 760px; margin-bottom: 34px; }
.application-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 34px; }
.application-card { border-top: 3px solid var(--gold); padding-top: 23px; min-width: 0; display: flex; flex-direction: column; scroll-margin-top: 30px; }
.application-card h3 { font-size: 27px; line-height: 1.25; }
.application-card h3 a { color: var(--ink); }
.application-card p { font-size: 15px; line-height: 1.8; }
.application-card .application-label { color: var(--accent); font-size: 12px; font-weight: 600; margin-bottom: 14px; }
.application-card .application-stage { margin-top: auto; margin-bottom: 0; padding-top: 16px; border-top: 1px solid var(--line); color: var(--muted); font-size: 11px; line-height: 1.65; }
.application-card .text-link { margin-top: 14px; font-size: 12px; }
.editing-preview { display: flex; align-items: center; justify-content: space-between; gap: 35px; background: var(--soft); padding: 30px 34px; margin-top: 38px; }
.editing-preview .eyebrow { margin-bottom: 10px; }
.editing-preview h3 { font-size: 24px; margin-bottom: 12px; }
.editing-preview p:last-child { font-size: 15px; max-width: 730px; margin: 0; }
.editing-preview > .text-link { flex-shrink: 0; }
.research-applications { margin-top: 40px; padding-top: 32px; border-top: 1px solid var(--line); scroll-margin-top: 30px; }
.research-applications > h2 { margin-top: 0; font-size: 33px; max-width: 700px; }
.research-applications .application-grid { margin-top: 32px; gap: 30px; }
.translation-note { border-block: 1px solid var(--line); padding-block: 24px; margin: 32px 0 0; font-size: 15px; }
.gene-editing-direction { display: grid; grid-template-columns: .85fr 1.4fr; gap: 42px; padding: 38px 34px; margin-top: 40px; background: var(--soft); border-top: 3px solid var(--gold); scroll-margin-top: 30px; }
.gene-editing-direction h2 { margin-top: 0; font-size: 33px; }
.gene-editing-direction p { font-size: 15px; }
.gene-editing-direction .research-status { font-size: 11px; }
.gene-editing-direction .editing-question { font-size: 19px; color: var(--ink); line-height: 1.6; }
.published-foundation-heading { padding-top: 58px; scroll-margin-top: 20px; }
.published-foundation-heading h2 { margin-top: 0; font-size: 33px; }
.published-study { display: grid; grid-template-columns: minmax(0, .85fr) minmax(0, 1.4fr); gap: 40px; align-items: start; padding-top: 42px; padding-bottom: 22px; }
.published-study::after { display: none; }
.published-study h2 { scroll-margin-top: 120px; }
.study-copy p { font-size: 15px; line-height: 1.8; }
.study-copy .research-status { font-size: 10px; line-height: 1.8; }
.study-copy .study-question { font-size: 18px; line-height: 1.5; font-weight: 600; color: var(--ink); }
.study-copy .study-outcome { border-left: 2px solid var(--accent); padding-left: 17px; color: var(--ink); }
.study-copy .research-keywords { font-size: 12px; }
.study-figure { margin: 0; min-width: 0; }
.study-image { display: block; padding: 14px; border: 1px solid var(--line); background: #fff; }
.study-image img { display: block; width: 100%; height: auto; max-height: 640px; object-fit: contain; }
.study-figure figcaption { padding: 16px 0 0; font-size: 12px; line-height: 1.75; color: var(--muted); }
.study-figure figcaption strong { color: var(--ink); }
.figure-credit { display: inline-block; margin-top: 6px; font-size: 11px; }
.figure-expand { display: block; width: fit-content; padding: 10px 0; font-size: 12px; font-weight: 600; }
.research-keywords { color: var(--accent); font-size: 13px; }
.research-status { color: var(--accent); font-size: 11px; font-weight: 650; text-transform: uppercase; letter-spacing: 1.2px; margin-bottom: 15px; }
.future-vision { margin-top: 44px; padding: 34px 36px; background: var(--soft); border-top: 3px solid var(--gold); scroll-margin-top: 25px; }
.future-vision h2 { margin-top: 0; font-size: 33px; }
.future-vision h3 { margin-top: 30px; font-size: 23px; letter-spacing: -.02em; }
.future-vision p:last-child { margin-bottom: 0; }
.callout { border-left: 3px solid var(--gold); background: var(--soft); padding: 24px 28px; margin-top: 32px; }
.callout p:last-child { margin-bottom: 0; }
.publication-summary { font-size: 14px; color: var(--muted); }
.publication-summary span { margin-inline: 8px; }
.publication-tools { margin: 25px 0 14px; }
.publication-tools > label { display: block; font-size: 13px; font-weight: 600; margin-bottom: 8px; }
.publication-tools input { background: var(--paper); color: var(--ink); border: 1px solid var(--line); width: 100%; min-height: 50px; padding: 12px 15px; border-radius: 0; font-size: 15px; }
.publication-tools input::placeholder { color: var(--muted); }
.filter-buttons { display: flex; gap: 10px; flex-wrap: wrap; margin: 14px 0; }
.filter-buttons button { min-height: 44px; font-size: 12px; border: 1px solid var(--line); color: var(--body); background: var(--paper); padding: 9px 14px; border-radius: 0; }
.filter-buttons button[aria-pressed="true"] { color: var(--paper); background: var(--ink); border-color: var(--ink); }
.publication-item { border-bottom: 1px solid var(--line); padding: 30px 0; }
.publication-meta { display: flex; align-items: baseline; justify-content: space-between; gap: 20px; font-size: 12px; text-transform: uppercase; letter-spacing: .9px; color: var(--accent); margin: 0 0 12px; }
.publication-meta > span:last-child { color: var(--muted); }
.publication-item h3 { font-size: 25px; letter-spacing: -.025em; font-weight: 600; line-height: 1.4; margin-bottom: 12px; }
.publication-item h3 a { color: var(--ink); }
.publication-authors { font-size: 14px; color: var(--muted); margin-bottom: 18px; }
.publication-authors strong { color: var(--ink); }
.publication-actions { display: flex; align-items: center; flex-wrap: wrap; gap: 22px; font-size: 12px; font-weight: 600; }
.badge { background: var(--soft); padding: 5px 8px; color: var(--accent); font-size: 11px; font-weight: 500; }
.small-note { font-size: 12px; color: var(--muted); margin-top: 20px; }
.book-entry { border-bottom: 1px solid var(--line); padding: 24px 0; }
.book-entry h3 { font-size: 23px; }
.book-entry p:not(.publication-meta) { font-size: 14px; color: var(--muted); }
.book-entry a { font-size: 13px; }
.book-entry h3 a { font: inherit; color: var(--ink); }
.mentoring-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 32px; border-block: 1px solid var(--line); padding-block: 32px; margin-block: 36px; }
.mentoring-grid h3 { font-size: 23px; }
.mentoring-grid p { font-size: 15px; margin-bottom: 0; }
.timeline-item { border-left: 1px solid var(--line); position: relative; padding: 0 0 22px 26px; }
.timeline-item::before { content: ''; position: absolute; width: 7px; height: 7px; border-radius: 50%; background: var(--accent); top: 10px; left: -4px; }
.timeline-date { color: var(--accent); font-size: 12px; letter-spacing: .6px; display: block; margin-bottom: 12px; }
.timeline-item h3 { font-size: 25px; margin-bottom: 12px; }
.timeline-item p { font-size: 16px; }
table { width: 100%; border-collapse: collapse; font-size: 15px; margin: 24px 0; }
th, td { border-bottom: 1px solid var(--line); text-align: left; padding: 14px 12px; vertical-align: top; }
th { background: var(--soft); font-size: 13px; color: var(--ink); font-weight: 600; }
.contact-card { border-top: 3px solid var(--gold); padding: 34px; background: var(--soft); margin-bottom: 30px; }
.contact-card h3 { font-size: 29px; }
.contact-card p { font-size: 17px; }
.contact-email { font-size: 21px; display: inline-flex; gap: 18px; max-width: 100%; overflow-wrap: anywhere; }
.publication-venue { color: var(--accent); font-size: 16px; }
.citation-text { background: var(--soft); padding: 25px; font-size: 14px; overflow-wrap: anywhere; }
.article-copy { margin: 28px 0; }
.filetype { font-size: 9px; border: 1px solid currentColor; padding: 2px 4px; }
.site-footer { color: #dce5ef; background: #092751; }
.site-footer a { color: inherit; }
.footer-main, .footer-bottom { width: min(1120px, 88%); margin: auto; display: flex; justify-content: space-between; gap: 30px; }
.footer-main { padding: 45px 0 37px; align-items: center; }
.footer-name { color: #fff !important; font-weight: 600; font-size: 24px; letter-spacing: -.5px; }
.footer-main p { font-size: 13px; line-height: 1.8; margin: 12px 0 0; }
.footer-contact { text-align: right; }
.footer-contact > a { font-size: 16px; }
.footer-contact > div { display: flex; gap: 24px; justify-content: flex-end; font-size: 12px; margin-top: 16px; }
.footer-bottom { border-top: 1px solid #345071; padding: 19px 0 23px; font-size: 10px; }
.footer-bottom p { margin: 0; }
.visually-hidden { position: absolute; width: 1px; height: 1px; padding: 0; margin: -1px; overflow: hidden; clip: rect(0,0,0,0); white-space: nowrap; border: 0; }
@media (max-width: 1100px) {
  .header-inner { gap: 20px; padding-inline: 4%; }
  .wordmark { font-size: 24px; }
  .wordmark-degree { display: none; }
  .wordmark-subtitle { font-size: 9px; letter-spacing: .2px; }
  .primary-links { gap: 20px; }
  .primary-links a { font-size: 16px; }
  .home-intro { gap: 42px; }
  .home-intro h2 { font-size: 33px; }
  .research-grid { gap: 28px; }
  .research-overview h3 { font-size: 21px; }
}
@media (max-width: 850px) {
  .application-grid, .gene-editing-direction { grid-template-columns: 1fr; }
  .application-grid { gap: 30px; }
  .application-card h3 br { display: none; }
  .application-card .application-stage { padding-top: 12px; }
  .editing-preview { align-items: flex-start; flex-direction: column; gap: 22px; }
  .gene-editing-direction { gap: 10px; }
  .home-applications { padding-top: 55px; }
  .published-study { grid-template-columns: 1fr; gap: 25px; }
  .study-copy p { font-size: 16px; }
  .study-figure { width: 100%; max-width: 650px; }
  .study-image { padding: 10px; }
  .header-inner { min-height: 92px; flex-wrap: wrap; padding: 22px 6%; gap: 0; }
  .wordmark { font-size: 25px; }
  .primary-links { width: 100%; order: 4; margin: 18px 0 0; justify-content: space-between; flex-wrap: wrap; gap: 14px; }
  .primary-links a { font-size: 16px; min-height: 44px; }
  .js .primary-links { display: none; }
  .js .primary-links.is-open { display: flex; }
  .js .menu-toggle { display: flex; align-items: center; gap: 8px; margin-left: auto; min-height: 44px; padding: 8px 9px; color: var(--ink); background: none; border: 0; font-size: 12px; }
  .appearance-toggle { margin-left: 12px; }
  .hero-inner { grid-template-columns: 1fr; width: min(580px, 88%); gap: 10px; padding: 10px 0 28px; }
  .hero-card { padding: 34px 0 14px; }
  .hero-card h1 { font-size: 50px; }
  .hero-tagline { font-size: 25px; }
  .hero-media { width: 100%; }
  .research-illustration { float: none; width: min(420px, 100%); margin: 24px 0; }
  .home-intro { grid-template-columns: 1fr; gap: 26px; padding-top: 48px; padding-bottom: 58px; }
  .profile-portrait { width: min(300px, 100%); }
  .home-intro h2 br { display: none; }
  .research-grid { grid-template-columns: 1fr; gap: 26px; }
  .research-overview { padding-top: 24px; }
  .research-overview h3 { font-size: 25px; }
  .research-overview h3 br { display: none; }
  .research-overview .direction-number { margin-bottom: 10px; }
  .research-arrow { position: absolute; top: 19px; right: 0; }
  .mentoring-grid { grid-template-columns: 1fr; }
  .footer-main { align-items: flex-start; }
}
@media (max-width: 520px) {
  .home-applications .section-heading h2, .research-applications > h2, .gene-editing-direction h2, .published-foundation-heading h2 { font-size: 29px; }
  .editing-preview, .gene-editing-direction { padding: 26px 22px; }
  body { font-size: 16px; }
  .wordmark { font-size: 23px; }
  .wordmark-subtitle { font-size: 8px; letter-spacing: .1px; }
  .appearance-toggle { margin-left: 3px; width: 44px; height: 44px; }
  .header-inner { padding: 18px 5%; min-height: 88px; }
  .js .menu-toggle { padding-inline: 5px; gap: 6px; }
  .hero-inner { width: 88%; gap: 18px; }
  .image-caption { font-size: 9px; text-align: left; padding-top: 10px; }
  .hero-card { padding: 30px 0 0; }
  .hero-card .eyebrow { font-size: 8px; letter-spacing: 1.1px; margin-bottom: 17px; }
  .hero-card h1 { font-size: 39px; line-height: 1.25; margin-bottom: 18px; }
  .hero-tagline { font-size: 20px; margin-bottom: 24px; }
  .hero-actions { gap: 16px; }
  .hero-actions .button { padding-inline: 16px; gap: 16px; font-size: 12px; }
  .hero-about { font-size: 12px; }
  .content-width, .main-content, .page-banner-inner { width: 88%; }
  .home-intro { padding-top: 43px; padding-bottom: 53px; }
  .home-intro h2, h2 { font-size: 30px; }
  .intro-copy p { font-size: 16px; }
  .home-research { padding: 48px 0 44px; }
  .section-heading { align-items: flex-start; flex-direction: column; gap: 17px; margin-bottom: 28px; }
  .section-heading h2 { font-size: 30px; }
  .research-overview h3 { font-size: 24px; }
  .home-news { padding-top: 50px; padding-bottom: 52px; }
  .update-row { grid-template-columns: 1fr; gap: 11px; padding: 22px 0; }
  .update-row p { font-size: 15px; }
  .update-row time { font-size: 10px; }
  .page-banner-inner { padding: 39px 0 35px; }
  .page-banner h1 { font-size: 36px; overflow-wrap: break-word; }
  .page-banner .eyebrow { font-size: 10px; letter-spacing: 1.3px; }
  .section-intro { font-size: 18px; }
  .main-content { padding: 36px 0 56px; }
  .research-direction h2 { font-size: 27px; }
  .research-keywords { font-size: 12px; }
  .future-vision { padding: 26px 22px; }
  .future-vision h2 { font-size: 28px; }
  .future-vision h3 { font-size: 22px; }
  .publication-meta { font-size: 10px; gap: 10px; }
  .publication-item h3 { font-size: 22px; }
  .publication-authors { font-size: 13px; }
  .publication-summary { font-size: 12px; }
  .publication-tools input { font-size: 13px; }
  .filter-buttons { gap: 7px; }
  .filter-buttons button { font-size: 10px; padding-inline: 10px; }
  .publication-actions { gap: 15px; }
  .contact-card { padding: 26px 23px; }
  .contact-email { font-size: 16px; gap: 8px; }
  table { font-size: 12px; }
  th, td { padding: 11px 6px; }
  .footer-main { flex-direction: column; gap: 30px; padding: 35px 0; }
  .footer-contact { text-align: left; }
  .footer-contact > div { justify-content: flex-start; }
  .footer-bottom { flex-direction: column; gap: 8px; }
}
@media (max-width: 360px) {
  .hero-card h1 { font-size: 33px; }
  .wordmark { font-size: 20px; }
  .wordmark-subtitle { font-size: 7px; }
}
@media (prefers-reduced-motion: reduce) { html { scroll-behavior: auto; } * { animation: none !important; transition: none !important; } }
@media print {
  .site-header, .site-footer, .publication-tools, .button, .hero-media { display: none !important; }
  body { color: #000; background: #fff; font-size: 11pt; }
  .home-hero { background: #fff; color: #000; }
  .hero-inner { display: block; width: 100%; padding: 0; }
  .hero-card { margin: 0; padding: 0; }
  .hero-card h1, .hero-card .eyebrow, .hero-tagline { color: #000; }
  .hero-actions { display: none; }
  .main-content, .content-width, .page-banner-inner { width: 100%; padding: 20px 0; }
  h1 { font-size: 28pt; } h2 { font-size: 20pt; }
  .publication-item, .research-direction { break-inside: avoid; }
}

/* Candidate homepage: image banner and an overlapping white introduction. */
.candidate-hero { padding-bottom: 50px; }
.candidate-banner { position: relative; isolation: isolate; background: #081c35; }
.candidate-banner::before { content: ''; position: absolute; z-index: -1; inset: 0 0 0 auto; width: min(850px, 80%); background: linear-gradient(90deg, #081c35, rgba(8,28,53,.55) 38%, rgba(8,28,53,.08)), url('../../images/rna-delivery-illustration.png') center 46% / cover no-repeat; }
.candidate-banner > div { min-height: 455px; padding-block: 86px 145px; }
.candidate-banner .eyebrow { color: #d2e1ee; font-size: 15px; margin-bottom: 21px; }
.candidate-banner h1 { color: #fff; font-size: clamp(40px, 4.5vw, 60px); line-height: 1.16; max-width: 710px; margin: 0; }
.candidate-banner h1 em { font-style: normal; color: #fff; }
.candidate-hero-inner { display: grid; grid-template-columns: 230px minmax(0, 1fr); gap: 58px; align-items: center; position: relative; margin-top: -70px; padding: 44px 58px 0; background: var(--paper); }
.candidate-name { color: var(--ink); font-size: 38px; font-weight: 700; margin-bottom: 9px; letter-spacing: -.8px; line-height: 1.2; }
.candidate-role { font-size: 15px; line-height: 1.7; color: var(--muted); margin-bottom: 20px; }
.candidate-lead { font-size: 20px; line-height: 1.65; max-width: 660px; margin-bottom: 24px; }
.hero-profile-links { display: flex; gap: 25px; flex-wrap: wrap; margin-top: 22px; font-size: 13px; }
.candidate-portrait { margin: 0; min-width: 0; }
.portrait-mat { background: #fff; }
.portrait-mat img { display: block; width: 100%; height: auto; }
.candidate-portrait figcaption { display: flex; flex-direction: column; gap: 8px; font-size: 11px; line-height: 1.5; margin-top: 15px; color: var(--muted); }
.candidate-portrait figcaption a { font-weight: 600; padding-block: 5px; }
.portrait-keywords { display: flex; flex-wrap: wrap; gap: 6px 12px; list-style: none; padding: 0 0 12px; margin: 0 0 4px; border-bottom: 1px solid var(--line); color: var(--ink); font-size: 13px; font-weight: 500; line-height: 1.6; }
.portrait-keywords li { margin: 0; }
.candidate-evidence { display: grid; grid-template-columns: 1fr 1.15fr 1.1fr; border-block: 1px solid var(--line); padding: 30px 0; gap: 30px; }
.candidate-evidence > div + div { border-left: 1px solid var(--line); padding-left: 30px; }
.evidence-label { display: block; color: var(--accent); font-size: 12px; font-weight: 600; margin-bottom: 10px; }
.candidate-evidence p { color: var(--ink); font-size: 16px; line-height: 1.5; margin: 0 0 6px; }
.candidate-evidence p strong { font-size: 25px; font-weight: 700; margin-right: 4px; }
.candidate-evidence > div > span:last-child { display: block; font-size: 12px; line-height: 1.6; color: var(--muted); }
.candidate-about { display: grid; grid-template-columns: 1fr 1.55fr; gap: 70px; padding-block: 90px; scroll-margin-top: 35px; }
.candidate-about h2 { font-size: 34px; margin: 0; line-height: 1.23; }
.about-narrative p { font-size: 17px; }
.about-narrative .about-lead { font-size: 22px; line-height: 1.55; color: var(--ink); }
.research-story-band { background: var(--soft); }
.research-story-inner { display: grid; grid-template-columns: 1fr 1.15fr; gap: 65px; align-items: center; padding-block: 74px; }
.research-story-art { margin: 0; }
.research-story-art img { width: 100%; height: auto; display: block; }
.research-story-art figcaption { font-size: 11px; line-height: 1.6; color: var(--muted); margin-top: 14px; }
.research-movie { display: block; width: 100%; height: auto; background: #000; }
.research-movie:focus-visible { outline: 3px solid var(--gold); outline-offset: 5px; }
.movie-title { display: block; font-size: 17px; color: var(--ink); margin-bottom: 8px; }
.movie-description { display: block; font-size: 13px; line-height: 1.7; color: var(--body); }
.movie-credit { display: block; margin-top: 10px; }
.research-story-copy h2 { font-size: 34px; line-height: 1.23; margin: 0 0 23px; }
.research-story-copy > p:not(.eyebrow) { font-size: 17px; line-height: 1.75; }
.barrier-list { list-style: none; margin: 27px 0; padding: 0; }
.barrier-list li { border-top: 1px solid var(--line); padding: 17px 0 0; display: flex; gap: 20px; margin: 0 0 17px; }
.barrier-list li > span { font-size: 13px; color: var(--accent); padding-top: 1px; }
.barrier-list h3 { font-size: 18px; letter-spacing: -.2px; margin: 0 0 6px; }
.barrier-list p { font-size: 14px; line-height: 1.7; margin: 0; }
.featured-research { padding-block: 90px; }
.featured-research .section-heading h2 { font-size: 36px; }
.featured-paper-grid { display: grid; grid-template-columns: repeat(3, minmax(0, 1fr)); gap: 38px; }
.featured-paper { display: flex; flex-direction: column; min-width: 0; }
.featured-paper-image { display: flex; align-items: center; justify-content: center; height: 260px; padding: 10px; background: #fff; border-bottom: 1px solid var(--line); }
.featured-paper-image img { display: block; width: 100%; height: 100%; object-fit: contain; }
.featured-paper-copy { padding: 25px 0 0; display: flex; flex-direction: column; flex: 1; }
.paper-kicker { font-size: 11px; color: var(--accent); display: flex; justify-content: space-between; gap: 8px; margin-bottom: 15px; font-weight: 600; }
.paper-kicker span { color: var(--muted); }
.featured-paper h3 { font-size: 25px; line-height: 1.25; font-weight: 700; letter-spacing: -.5px; margin-bottom: 17px; }
.featured-paper h3 a { color: var(--ink); }
.featured-paper-copy > p:not([class]) { font-size: 15px; line-height: 1.75; }
.paper-role { color: var(--accent); font-size: 12px; font-weight: 600; margin: auto 0 17px; }
.paper-links { display: flex; gap: 24px; font-size: 13px; font-weight: 600; padding-top: 15px; border-top: 1px solid var(--line); }
.paper-credit { font-size: 10px; color: var(--muted); margin: 17px 0 0; line-height: 1.6; }
.candidate-next { background: var(--soft); }
.candidate-next-inner { display: grid; grid-template-columns: 1fr 1fr; gap: 75px; padding-block: 76px; }
.candidate-next h2 { font-size: 32px; line-height: 1.25; margin: 0 0 23px; }
.candidate-next p:not(.eyebrow) { font-size: 17px; line-height: 1.75; }
.next-mentoring { border-left: 1px solid var(--line); padding-left: 50px; }
.candidate-updates { padding-block: 80px 40px; }
.candidate-updates h2 { font-size: 36px; }
.candidate-contact { display: flex; justify-content: space-between; align-items: center; gap: 40px; padding-block: 44px 80px; }
.candidate-contact h2 { font-size: 34px; margin: 0; }
.candidate-contact .button { flex-shrink: 0; }
@media (max-width: 1100px) {
  .candidate-hero-inner { gap: 38px; padding-inline: 38px; }
  .candidate-about { gap: 42px; }
  .research-story-inner { gap: 40px; }
  .featured-paper-grid { gap: 26px; }
  .featured-paper h3 { font-size: 23px; }
  .paper-kicker { flex-wrap: wrap; }
}
@media (max-width: 850px) {
  .candidate-banner > div { min-height: 400px; padding-block: 68px 115px; }
  .candidate-banner::before { width: 90%; }
  .candidate-banner h1 { font-size: 46px; }
  .candidate-hero-inner { grid-template-columns: 180px 1fr; gap: 30px; padding: 32px 28px 0; margin-top: -50px; }
  .candidate-name { font-size: 31px; }
  .candidate-lead { font-size: 18px; }
  .candidate-evidence { gap: 20px; }
  .candidate-evidence > div + div { padding-left: 20px; }
  .candidate-about { grid-template-columns: 1fr; gap: 28px; padding-block: 62px; }
  .candidate-about h2 br { display: none; }
  .research-story-inner { grid-template-columns: 1fr; gap: 36px; padding-block: 55px; }
  .research-story-art { width: min(480px, 100%); margin: auto; }
  .research-story-copy { max-width: 640px; }
  .featured-paper-grid { grid-template-columns: 1fr; gap: 36px; }
  .featured-paper { display: grid; grid-template-columns: 1fr 1.2fr; gap: 28px; padding-top: 20px; border-top: 1px solid var(--line); }
  .featured-paper-image { height: 100%; min-height: 285px; border: 0; }
  .featured-paper-image img { max-height: 310px; }
  .featured-paper-copy { padding: 0; }
  .featured-paper h3 br { display: none; }
  .candidate-next-inner { gap: 35px; }
  .next-mentoring { padding-left: 35px; }
  .candidate-next h2 { font-size: 28px; }
}
@media (max-width: 600px) {
  .candidate-banner::before { width: 100%; background-position: center, center 46%; }
  .candidate-banner > div { min-height: 355px; padding-block: 55px 90px; }
  .candidate-banner h1 { font-size: clamp(33px, 8vw, 45px); line-height: 1.2; }
  .candidate-banner .eyebrow { font-size: 12px; }
  .candidate-hero { padding-bottom: 35px; }
  .candidate-hero-inner { grid-template-columns: 1fr; gap: 28px; padding: 28px 24px 0; margin-top: -38px; }
  .candidate-portrait { width: 165px; justify-self: center; }
  .candidate-portrait figcaption { font-size: 10px; }
  .candidate-name { font-size: 30px; }
  .candidate-role { font-size: 14px; }
  .candidate-lead { font-size: 18px; }
  .hero-profile-links { gap: 20px; }
  .candidate-evidence { grid-template-columns: 1fr; padding-block: 25px; gap: 19px; }
  .candidate-evidence > div + div { padding: 18px 0 0; border-left: 0; border-top: 1px solid var(--line); }
  .candidate-evidence .evidence-label { margin-bottom: 6px; }
  .candidate-about { padding-block: 52px; }
  .candidate-about h2, .featured-research .section-heading h2, .research-story-copy h2 { font-size: 30px; }
  .about-narrative .about-lead { font-size: 20px; }
  .research-story-copy h2 br { display: none; }
  .featured-research { padding-block: 55px; }
  .featured-paper { display: flex; gap: 22px; }
  .featured-paper-image { height: 265px; min-height: 0; }
  .featured-paper h3 { font-size: 26px; }
  .candidate-next-inner { grid-template-columns: 1fr; padding-block: 52px; }
  .next-mentoring { padding: 33px 0 0; border-left: 0; border-top: 1px solid var(--line); }
  .candidate-next h2 { font-size: 29px; }
  .candidate-next h2 br { display: none; }
  .candidate-updates { padding-block: 52px 20px; }
  .candidate-updates h2 { font-size: 30px; }
  .candidate-contact { flex-direction: column; align-items: flex-start; padding-block: 34px 54px; gap: 25px; }
  .candidate-contact h2 { font-size: 29px; }
  .candidate-contact h2 br { display: none; }
}
@media print {
  .candidate-hero-inner, .candidate-about, .candidate-next-inner { display: block; }
  .candidate-portrait, .research-story-art, .featured-paper-image, .hero-profile-links { display: none; }
  .candidate-hero, .candidate-about, .featured-research, .candidate-next-inner { padding-block: 20px; }
  .candidate-banner { background: none; }
  .candidate-banner::before { display: none; }
  .candidate-banner > div { min-height: 0; padding: 20px 0; }
  .candidate-banner h1, .candidate-banner h1 em, .candidate-banner .eyebrow { color: #000; }
  .candidate-hero-inner { margin-top: 0; padding: 20px 0; }
  .research-story-band { background: #fff; color: #000; }
  .research-story-copy h2, .barrier-list h3 { color: #000; }
  .next-mentoring { border: 0; padding: 20px 0 0; }
  .candidate-evidence { break-inside: avoid; }
}